Phalaenopsis Journal Third Quarter 2013 | Page 20

Understanding Phalaenopsis Flower Color Genetics Rob Griesbach Figure 1. A Phalaenopsis bellina cell containing chlorophyll Figure 2. Two cells showing the effects of an admixture of purple anthocyanin and within green chloroplasts and anthocyanin within the vacuole. yellow carotenoids leading to red color. “Genes act in mysterious ways, particular those with which we are not well acquainted.” I - Hugo Freed n Phalaenopsis, flower color is due to three different pigments – chlorophyll, flavonoids, and carotenoids (figure 1). Chlorophyll and carotenoids are both fatsoluble pigments. Green chlorophyll is found within the cell’s chloroplasts; while, yellow to orange carotenoids are found in chromoplasts. Flavonoids are water-soluble and found diffused within the cell’s large central vacuole. The flavonoids can be artificially subdivided into two groups, the anthocyanins and the co-pigments. The anthocyanins are red to blue in color; while, the co-pigments are cream colored (Griesbach, 2005). By mixing these pigments, different colors can be created. For example, a flower may appear red for several reasons: 1) the presence of a red anthocyanin and the absence of chlorophyll and carotenoids or 2) the presence of a purple anthocyanin combined with an orange carotenoid (figure 2). In Phalaenopsis the bars are red due to a purple anthocyanin and an orange carotenoid. The bars within the center of Phalaenopsis mariae flowers appear purple because the accompanying orange carotenoids are absent (figure 3). Most red Phalaenopsis hybrids are 20 Phalaenopsis | Third Quarter | Volume 23 | 2013 the result this ‘admixture’ of orange carotenoids and purple anthocyanins (Griesbach, 1984). Flavonoids are characterized by their unique 3 ring chemical skeleton. In nature, the flavonoids are found with one of more sugars and organic acids attached to the skeleton. The anthocyanins are the class of flavonoids that have hydroxyl groups (-OH) at the 3, 5 and 7 positions on the skeleton. Specific anthocyanins are dis- Figure 3. Phalaenopsis mariae ‘Zuma Canyon’ showing the difference in color of bars due to admixture of carotenoid and anthocyanin pigments.?