NEXT GENERATION GENOME EDITING TECHNOLOGIES Next-Generation-Genome-Editing-Technologies | Page 11

ALTERNATIVE SOLUTIONS: With the ongoing strife for rights to CRISPR acquisition, other researchers who have keen interest in gene editing tools or who need developing a much better tool than CAS9 which could overcome the problems related to it have laid their eyes on two new emerging tools: 1. Cpf1 The history of gene editing became interesting when a new nuclease came into picture. Zhang’s lab recently published a paper describing two genes from Cpf1 family that have cleaving ability in mammalian cells. 1. It is a class II nuclease (CRISPR from Prevotella and Francisella) and is classified as type V CRISPR system. 2. It has RuvC-like endonuclease which is similar to CAS9’s but lacks in HNH domain. 3. It requires only one RNA whereas CAS9 required two (trRNA and crRNA). 4. It cleaves DNA in a staggered manner and gives 5’overhang 18-20 bp away from PAM sites. CAS9 produces blunt ended double strands (3nt upstream of PAM sites). 5. The Cpf1 crRNA has a much simpler structure (short stem loop in direct repeat). Benefits: ƒ ƒ Due to staggered cleavage, directional gene transfer is possible ƒ ƒ Sticky end mediated gene transfer can be used to target nondividing cells, which are difficult to modify through HDR. ƒ ƒ AT- rich regions that did not have enough 3’-NGG PAM sites can also be treated as targets.