Maximum Yield USA 2015 December | Page 56

POWERING PHOTOSYNTHESIS With the stomata shut to prevent water loss, photosynthesis cannot occur, as CO2 can’t be taken in from the surrounding air. Plant growth and yield will be slowed if this occurs too often. When humidity is low, particularly when combined with warm growing conditions, it can be difficult to tell if plants have shut down photosynthesis due to closed stomata, as under these conditions, the foliage may look normal. One way of checking to see if the stomata are open is to measure the difference between leaf surface and air temperature using a handheld infrared thermometer. A plant with open stomata is actively cooling itself via transpiration, so the leaf surface will be a slightly lower temperature than the surrounding air. If the leaf surface is warmer than the surrounding air, then transpiration is not occurring and photosynthesis has shut down. An infrared thermometer can be used to determine leaf temperature and whether the stomata are open for CO2 uptake. Along with light quality and quantity and CO2 levels, temperature also determines the rate of glucose production during photosynthesis.” Tomato plants respond well to CO2 supplementation. Light and Temperature Low-light symptoms include plant elongation and stretching. 54 Maximum Yield USA  |  December 2015 Along with light quality and quantity and CO2 levels, temperature also determines the rate of glucose production during photosynthesis due to the activity of enzymes that catalyze biochemical reactions in leaf tissues. These enzymes are strongly affected by temperature, so the rate of assimilate production through the process of photosynthesis is maximized when the plant is grown in its optimal temperature range. At low temperatures, which for many plants we commonly grow is 32-50°F, the enzymes involved in photosynthesis do not work at full efficiency and this lowers the rate of sugar production and plant yields. At optimal temperatures, which vary for different species, but are often between 50 and 75°F, photosynthetic enzymes work at their maximum rate. At these temperatures, it is often the diffusion of CO2 that becomes the limiting factor for photosynthesis. At high temperatures (104-113°F), many plant species will experience a drop in photosynthetic rate.