iHerp Australia Issue 1 | Page 5

A mphibians are the most endangered group of vertebrates on the planet. The IUCN (International Union for the Conservation of Nature) estimates that about 33% of the 7,000 or so recognised species are threatened with extinction. Over the past three decades amphibians have been disap- pearing at an alarming rate. Over 200 species have become extinct, and about 43% of all amphibians are considered to be declining – and this is likely to be an underestimate given how little is known about many of these unassuming animals and their population dynamics. Indeed, new species of amphibians continue to be discovered every year, many of which are vulner- able because they are restricted to small geographic ranges. Moreover, for at least a quarter of the species currently recognised, there is insufficient available data to even form an assessment of their conservation status. The imperilled state of amphibians is not confined to poor regions overseas: in Australia almost one quarter of species are acknowl- edged to be of conservation concern. The threats faced by amphibians are numerous: habitat loss and degradation (largely stemming from land clearing to make way for livestock); climate change (driven by massive use of fossil fuels, with the livestock industry also being a key contributor); collection for consumption and the pet trade; pollution including pesticides; and disease. All except the latter are attributable to the detrimental effects human- kind’s exploitive actions have upon the environment; hence, in theory, they can be readily corrected and prevented. Disease is a lot more difficult to address, yet disease, often acting in addition to or in synergy with other factors, is one of the major reasons amphibians across the globe are in jeopardy. Chytridiomycosis is an emerging infectious disease caused by the aquatic fungal pathogen, Batrachochytrium dendrobatidis (or Bd), a chytrid fungus. Bd infects the superficial, keratin- containing layers of amphibian skin, and the mouthparts of tadpoles. This inhibits osmoregu- lation, ionoregulation and cutaneous respiration. Symptoms include lethargy, sitting in an abnormal position and in the sun, excessive skin sloughing, and eventually death. ‘ Bd and in protected areas. Against the global amphibian extinction crisis and ongoing catastrophic declines of amphibi- ans in the face of Bd, there is nevertheless some hope. One amazing Australian species has been confirmed to have begun the road to recovery: Fleay’s Barred Frog (Mixophyes fleayi). The effects of Bd have been particularly prevalent amongst native stream-breeding frogs like is responsible for the greatest loss of biodiversity disease in due to recorded history history.’ Bd is responsible for the greatest loss of biodiversity due to disease in recorded history, and more than 350 amphibian species are known to have been infected. Australia’s anuran fauna have been hard hit, with rapid declines beginning in the late 1970s in southern Queensland, near Brisbane. In fact, it was Australian scientists who, in 1996, first posited that disease was the culprit behind the devas- tation of frog populations. The pathogen responsible was confirmed in 1998, and Bd was described in 1999. Since then, exhaustive efforts have been made to elucidate the genetics and biology of the fungus and the disease it causes, and to devise strategies to prevent its potentially cataclysmic impact. Despite some progress, much remains unknown, and there is no universal cure for chytridiomy- cosis or means of eradicating the pathogen that causes it. Many proven strategies for species preservation - like habitat protec- tion - are insufficient to stem declines caused by this disease, which has wiped out populations in the most pristine of habitats M. fleayi, with over ten species from the subtropical and tropical rainforests of eastern Australia suffering serious declines since 1979. In this same region Bd has also caused the extinction of four species: the Southern Gastric- brooding Frog (Rheobatrachus silus - last seen 1981); the Northern Gastric-brooding Frog (R. vitellinus – 1985); the Sharp- snouted Day Frog (Taudactylus acutirostris - 1997); and the Southern Day Frog (T. diurnus - 1979). In southeastern Queensland and adjacent New South Wales, near where declines from Bd were first detected in the 1970s, popula- tions of M. fleayi, along with the closely-related Giant Barred Frog (Mixophyes iteratus), and the Cascade Tree Frog (Litoria pearsoniana), underwent precipi- tous collapses over just a few short years. With declines of over 90%, it seemed these species were also heading inexorably towards extinction. Mixophyes fleayi has a narrow distribution along the Great Dividing Range and coast from the Conondale Range in south- east Queensland to the vicinity of