Gazelle : The Palestinian Biological Bulletin (ISSN 0178 – 6288) . Number 120, December 2014, pp. 1-29. | Page 16

16 The skeleton of the Long-necked Whip-tailed Sauropod Dinosaur (Amphicoelias brontodiplodocus Galiano and Albersdörfer, 2010) at Dubai Mall, Dubai, UAE. Photo by Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. 12.03.2014. https://www.flickr.com/photos/50022881@N00/13963351962/ Throughout their evolutionary history, sauropod dinosaurs were found primarily in semi-arid, seasonally dry environments, with a corresponding seasonal drop in the quality of food during the dry season. The environment of Amphicoelias was essentially a savanna, similar to the arid environments in which modern giant herbivores are found, supporting the idea that poor-quality food in an arid environment promotes the evolution of giant herbivores. Carpenter argued that other benefits of large size, such as relative immunity from predators, lower energy expenditure, and longer life span, are probably secondary advantages (Wikipedia). The Morrison Formation environment in which Amphicoelias lived would have resembled a modern savanna, though since grass did not appear until the Late Cretaceous, ferns were probably the dominant plant and main food source for Amphicoelias. Though Engelmann et al. (2004) dismissed ferns as a sauropod food source due to their relatively low caloric content; Carpenter argued that the sauropod digestive system, well adapted to handle low-quality food, allows for the consumption of ferns as a large part of the sauropod diet. Carpenter also Gazelle : The Palestinian Biological Bulletin – Number 120 – December 2014