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INVESTIGATIVE REPORT
Antipruritic Effects of Janus Kinase Inhibitor Tofacitinib in a
Mouse Model of Psoriasis
Takashi HASHIMOTO, Kent SAKAI, Kristen M. SANDERS, Gil YOSIPOVITCH and Tasuku AKIYAMA
Department of Dermatology and Cutaneous Surgery, University of Miami, Miami Itch Center, Miami, FL, USA
The Janus kinase 1/3 inhibitor tofacitinib has demon-
strated an antipruritic effect in two phase ΙΙΙ studies
in psoriasis. However, the mechanisms behind this
antipruritic effect are still unknown. We presently in-
vestigated whether tofacitinib affects spontaneous
itch as well as expression of itch-related cytokines and
epidermal nerve fiber density (ENFD) in the imiqui-
mod-induced mouse model of psoriasis. Psoriasis-like
skin lesions were produced by daily topical application
of imiquimod to the back skin. Imiquimod treatment
resulted in spontaneous scratching, which was signi-
ficantly inhibited by tofacitinib treatment. Imiquimod
treatment significantly increased mRNA expression of
Il22, Il23, and Il31, reduced peptidergic ENFD, and in-
creased nonpeptidergic ENFD compared to naive mice.
Tofacitinib significantly decreased the expression of
those cytokines and increased peptidergic ENFD with
out a significant effect on nonpeptidergic ENFD. Tofa-
citinib may inhibit psoriatic itch through inhibition of
cytokine expression as well as modulation of epider-
mal innervation.
Key words: JAK inhibitor; chronic itch; psoriasis; scratching;
IL-31.
Accepted Nov 20, 2018; E-published Nov 21, 2018
Acta Derm Venereol 2019; 99: 298–303.
Corr: Tasuku Akiyama, PhD, Department of Dermatology and Cuta-
neous Surgery, University of Miami, Miami Itch Center, 1600 NW 10 th Ave
RMSB2063, Miami, FL 33136, USA. E-mail: [email protected]
T
ofacitinib is a targeted small molecule that preferen-
tially inhibits Janus kinase (JAK)-1/3. This inhibition
of JAK-1/3 blocks γ-chain cytokines, including inter-
leukin (IL)-2, IL-4, IL-7, IL-9, IL-15, and IL-21, that
could contribute to skin inflammation (1). Tofacitinib
demonstrated a significant antipruritic effect in two phase
ΙΙΙ studies of patients with moderate to severe plaque
psoriasis and one phase ΙΙa study of patients with mild to
moderate atopic dermatitis (2–4). Interestingly, tofaciti-
nib has a rapid antipruritic effect which is independent
from improvements in other clinical signs of psoriasis,
such as erythema, induration, and scaling (5). A recent
study suggests that tofacitinib acts directly on the trans-
ient receptor potential vanilloid 1 ion channel expressed
in sensory neurons to exert antipruritic effects (6). This
could explain the rapid onset of improvement in pruritus.
However, the mechanism behind the antipruritic effects
of tofacitinib has not been fully understood.
doi: 10.2340/00015555-3086
Acta Derm Venereol 2019; 99: 298–303
SIGNIFICANCE
Tofacitinib, a drug that inhibits Janus kinases 1 and 3, has
been shown to reduce itch in clinical trials of psoriasis.
However, the exact mechanism for this anti-itch effect is
still unclear. This study investigates the effects of tofaciti-
nib treatment in a mouse model of psoriasis. Tofacitinib
reduced scratching in this model. Additionally, tofacitinib
rescued the increased mRNA levels of Il22, Il23, and Il31,
3 cytokines that are known to be related to itch. Finally,
tofacitinib recovered the reduction in peptidergic nerves in
the epidermis of psoriasis model mice. Therefore, tofaciti-
nib could reduce itch through regulation of cytokines and
epidermal nerves.
Psoriasis is an inflammatory skin disease that is driven
by Th1 and Th17 cells (7). IL-17A, IL-17F, IL-21, and
IL-22 are found at increased levels in psoriatic skin (8,
9). Interestingly, RNA-seq analysis demonstrated that
mRNA transcripts for Il17a, Il22, Il23, and Il31 were
elevated in the itchy skin of both atopic dermatitis and
psoriasis patients compared to non-itchy skin (10).
Knockout mouse studies have revealed the role of cyto-
kines (e.g. IL-31, thymic stromal lymphopoietin (TSLP),
IL-4, IL-33, and tumor necrosis factor-α) in itch (11–15).
Particularly, IL-31 and TSLP directly act on their cognate
cytokine receptors expressed by primary sensory neurons
and keratinocytes, respectively, to elicit itch.
Dynamic changes in epidermal innervation have
been observed under chronic itch conditions. Epidermal
hyperinnervation by itch-signaling fibers is considered
to be one cause of chronic itch (16), and increased epi-
dermal nerve density has been frequently reported in
humans with chronic itch conditions including psoriasis
(16–20). In line with this, intraepidermal nonpeptidergic
nerve density is increased in animal models of dry skin
itch and psoriatic itch (21–23). In contrast, conflicting
results have been reported for epidermal nerve density
in human with chronic itch. Reduced epidermal nerve
density has been reported in patients with psoriatic itch
as well as prurigo nodularis (17, 24, 25). Additionally,
intraepidermal peptidergic nerve density is decreased
in animal models of psoriatic itch (23). In these cases,
reduced primary afferent drive toward itch-inhibitory
interneurons could contribute to disinhibition of itch.
We presently investigated whether tofacitinib inhibited
spontaneous itch in the imiquimod-induced mouse model
This is an open access article under the CC BY-NC license. www.medicaljournals.se/acta
Journal Compilation © 2019 Acta Dermato-Venereologica.