Acta Dermato-Venereologica 99-13CompleteContent | Page 30
SHORT COMMUNICATION
1303
rs34567942 a Novel Susceptibility Single Nucleotide Polymorphism for Cutaneous Squamous Cell
Carcinoma in Organ Transplant Recipients
Aleksandar KUZMANOV 1 , Weihong QI 2 , Nadja STENZ 3 , Pierre-Yves BOCHUD 4 , Zoltan KUTALIK 5,6 , Agnieszka WÓJTOWICZ 7 ,
Gunther HOFBAUER 1 and the Swiss Transplant Cohort Study
1
Department of Dermatology, University Hospital Zurich, Wagistrasse 14, CH-8952 Schlieren, 2 Functional Genomics Centre Zurich, University
of Zürich, 3 Medical Faculty, University of Zürich, Zürich, 4 Service of Infectious Diseases, University Hospital and University of Lausanne,
5
Institute of Social and Preventive Medicine, University Hospital Lausanne (CHUV), 6 Swiss Institute of Bioinformatics, and 7 Service of Infectious
Diseases, University Hospital and University of Lausanne, Lausanne, Switzerland. E-mail: [email protected]
Accepted Sep 25, 2019; E-published Sep 25, 2019
Cutaneous squamous cell carcinoma (cSCC) is the se-
cond most common type of solid human tumour and a
main cause of cancer-related death in the general popu-
lation (1, 2). Typically, it emerges on ultraviolet (UV)-
exposed sun-damaged skin from benign intraepithelial
lesions called actinic keratosis (AK) (3, 4). High-risk
patient groups, such as organ transplant recipients (OTR),
display a 65–250-fold increase in cSCC, making cSCC
the most frequent cancer in this group (5). However, not
all OTR develop cSCC, while some develop a multi
plicity of sSCC (1). This discrepancy is the major focus
of the current study.
Specific genetic factors, such as single nucleotide po-
lymorphisms (SNPs) determining cSCC susceptibility in
OTRs, have been little-studied. Only a few papers have
addressed this topic (6–8). Moreover, only one group has
performed genome-wide association studies (GWAS),
replicating 10 candidate SNPs previously associated with
skin cancer in the general population; however, they did
not show SNP-significant genome-wide association with
cSCC in the OTRs (7). Our GWAS revealed novel OTR-
specific SNP associated with cSCC susceptibility in OTR.
42,400,475. Exclusion criteria were: violation of Hardy-Weinberg-
principle (p<1 E-6 ), call rate < 0.95, minor allele frequency (MAF)
< 0.01, and minor allele count (MAC) < 3. Untyped variants were
imputed using a combined reference panel of the 1,000 Genomes
Project phase 3 (9) and Genome of the Netherlands v5 (10) total-
ling more than 90 million genetic variants across the genome.
The software package SHAPEIT (11) was used for phasing and
IMPUTE2 (12) for imputation. The info statistic was computed
to establish imputation accuracy and markers with info < 0.7 were
excluded from further analysis.
For the complex MHC region, imputation of SNPs, multi-allelic
markers, amino acids, and classical HLA alleles using validated
SNP2HLA pipelines were performed. Preliminary results in a
subset of samples with HLA serology data showed that accuracy
is very high for HLA-A, -B, -DRB1, and DQB1 (> 95%), and high
for HLA-C (90%) in Europeans.
Manhattan plots were generated using the R package qqman
(http://biorxiv.org/content/biorxiv/early/2014/05/14/005165.full.
pdf). SNPs were queried against dbSNP (https://www.ncbi.nlm.
nih.gov/projects/SNP/) and the ensembl variation database (http://
www.ensembl.org/info/genome/variation/index.html). The results
were combined with annotation results obtained with snpEff (13).
GWAS identified 1 SNP, rs34567942, to be significantly as-
sociated with cSCC in OTRs at the p-value threshold of 5×10 –8
(Fig. 1). Table I shows the characteristics of SNP rs345567942;
namely the alternative nucleotide, imputation accuracy and im-
METHODS AND RESULTS
Patient data and material were collected prospectively from the
Swiss Transplant Cohort Study (STCS), from adult solid-organ
transplant recipients who received either kidney, liver, lung, heart,
pancreas, small bowel or mixed organ transplant between 2008 and
2011. Skin cancer episodes after or during transplantation were
reviewed by an independent clinician. All patients had provided
written informed consent for participation in the STCS (including
genetic analyses). The protocol was approved by the independent
ethics committees of each Swiss participating centre (University
Hospital of Lausanne; University Hospitals of Geneva; University
Hospital Zürich; Cantonal Hospital St Gallen; Inselspital, Bern
University Hospital; ClinicaLuganese, Lugano; and University
Hospital of Basel and registered at ClinicalTrials.gov Identifier
NCT01204944). To identify susceptibility SNPs for cSCC, GWAS
were performed on 61 OTR patients with cSCC and 908 skin
cancer negative-OTR patients. Genome-wide genotyping was
performed on DNA samples extracted from EDTA blood using
the Illumina ® Human Omni Express chip Genome Studio software
and default parameters were used to call genotypes. GWAS were
conducted using frequentist association tests in SNPtest. Asso-
ciations meeting the genome-wide threshold p-values of 5 × 10 –8
were considered statistically significant.
The raw genotype data-set included 719,665 variants. After ex-
clusion, 627,443 remained. The total number of imputed SNPs was
Fig. 1. Manhattan plot visualizing genome-wide association of
61 cutaneous squamous cell carcinoma (cSCC)-organ transplant
recipients (OTRs) and 908 skin cancer negative-OTRs, showing
–log10 p-value of SNPtest along chromosomes. Horizontal red line
represents the significant threshold p-value of 5×10 –8 . Arrow indicates single
nucleotide polymorphism rs34567942 with the most prominent p-value.
This is an open access article under the CC BY-NC license. www.medicaljournals.se/acta
Journal Compilation © 2019 Acta Dermato-Venereologica.
doi: 10.2340/00015555-3322
Acta Derm Venereol 2019; 99: 1303–1304