Acta Dermato-Venereologica 97-4 | Page 13
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INVESTIGATIVE REPORT
The Value of FLG Null Mutations in Predicting Treatment Response
in Atopic Dermatitis: An Observational Study in Finnish Patients
Tiia Maria LUUKKONEN 1,2# , Ville KIISKI 3# , Maria AHOLA 3 , Johanna MANDELIN 3 , Hannele VIRTANEN 3 , Minna PÖYHÖNEN 4,5 , Sirpa
KIVIRIKKO 4 , Ida SURAKKA 1,2 , Sakari REITAMO†, Aarno PALOTIE 1,6 , Markku HELIÖVAARA 2 , Eveliina JAKKULA 4 and Anita
REMITZ 3
Institute for Molecular Medicine Finland FIMM, 2 National Institute for Health and Welfare THL, 3 Skin and Allergy Hospital, 4 Clinical Genetics,
University of Helsinki and Helsinki University Hospital, 5 Department of Medical Genetics, Haartman Institute, University of Helsinki, Helsinki,
Finland, and 6 Broad Institute of Harvard and MIT, Cambridge, MA, USA. †Deceased
#
These authors contributed equally to this work.
1
The contribution of FLG null mutations to predicting
atopic dermatitis (AD) treatment response is not clear,
nor have such mutations been studied in the Finnish
population. This study tested the association of the
4 most prevalent European FLG null mutations, the 2
Finnish enriched FLG null mutations, the FLG 12-re-
peat allele, and 50 additional epidermal barrier gene
variants, with risk of AD, disease severity, clinical
features, risk of other atopic diseases, age of onset,
and treatment response in 501 patients with AD and
1,710 controls. AD, early-onset AD, palmar hyperlinea-
rity, and asthma showed significant associations with
the combined FLG null genotype. Disease severity and
treatment response were independent of patient FLG
status. Carrier frequencies of R501X, 2282del4, and
S3247X were notably lower in Finns compared with
reported frequencies in other populations. This data
confirms FLG mutations as risk factors for AD in Finns,
but also questions their feasibility as biomarkers in
predicting treatment response.
Key words: atopic dermatitis; filaggrin gene; skin barrier; tight
junction; biomarker; treatment response.
Accepted Nov 10, 2016; Epub ahead of print Nov 14, 2016
Acta Derm Venereol 2017; 97: 456–463.
Corr: Anita Remitz, Department of Dermatology, Allergology and Vene-
reology, Skin and Allergy Hospital, University of Helsinki and Helsinki Uni-
versity Hospital, PO Box PL 160, FIN-00029 Helsinki, Finland. E-mail:
[email protected]
A
topic dermatitis (AD; atopic eczema) is a chronic
or relapsing, itchy, inflammatory skin disease with
multifactorial pathogenesis and interplay of complex
underlying genetic and environmental factors. The skin
symptoms often prelude other manifestations of atopy:
allergic rhinitis, allergic conjunctivitis, and asthma (1).
The key role of epidermal barrier dysfunction in the
pathogenesis of AD is undisputable: the epidermis of AD
patients is characterized by a significant barrier impair-
ment, contributing to increased susceptibility to bacterial
and viral infections, microbial colonization, and allergic
sensitizations (2, 3).
Filaggrin is one of the most important proteins invol-
ved in epidermal barrier homeostasis. The filaggrin gene
(FLG) is a complex, highly polymorphic, and repetitive
doi: 10.2340/00015555-2578
Acta Derm Venereol 2017; 97: 456–463
gene located within the epidermal differentiation com-
plex on chromosome 1q21 (4). (The genomic and protein
organization of FLG is shown in Fig. 1). At least 3 FLG
copy number variants are recognized, with 10, 11 and
12 repeat alleles present in normal populations of Eu-
ropean ethnicity (4). Some studies have suggested that
extra filaggrin repeats, and thus extra filaggrin protein,
may strengthen the epidermal barrier properties of the
skin and consequently have a dose-dependent effect on
disease severity in AD (5, 6).
Prevalent FLG loss-of-function (LoF) mutations
(FLG null) are the most significant and consistently
replicated risk factors for AD in European (2282del4,
R501X, and R2447X) and Asian (3321delA and
Q2417X) populations (7–9). FLG null mutations have
been associated with earlier onset of disease (10). It has
been proposed that they are a risk factor only for the
early-onset form of AD (11) and they seem to increase
the risk of allergic rhinitis and food allergies in child-
hood and adolescence (12, 13). FLG null mutations are
also well-established risk factors for asthma in patients
with AD, but with regard to asthma in the absence of
eczema, results are conflicting (14, 15). FLG null muta-
tions have been associated with palmar hyperlinearity,
keratosis pilaris, hand eczema, and more persistent
symptoms of AD (9, 16–19).
The movement of substances between cells is regulated
by tight junctions (TJ), which form the major paracellular
barrier in the human epithelia (20). The permeability of
TJs is regulated mainly by claudins. Claudin 1 (CLDN1)
and Claudin 23 (CLDN23) expressions are reduced in
AD and the reduction in CLDN1 has been shown to di-
minish the integrity of TJs and correlates inversely with
Th2 cytokines (21, 22). Certain mutations of CLDN1 are
associated with increased risk of AD in North American
populations (21). The possible roles of many other TJ
proteins, and proteins involved in the structure and fun-
ction of the cell envelope structure in the pathogenesis
of AD are yet to be elucidated.
It seems plausible that AD consists of several different
endophenotypes, which may have an effect on response
to treatment (23). Studies on the effect of FLG null mu-
tations or other barrier gene mutations on the treatment
response are sparse.
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